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The 500 Dalton rule suggests small peptides can cross the stratum corneum, but intact copper coordination and hydrophilicity reduce net penetration

Olszewska-Pazdrak B, Hein TW, Olszewska P, Carney DH (2009) Chronic hypoxia attenuates VEGF signaling and angiogenic responses by downregulation of KDR in human endothelial cells

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R., Lower Activity of Superoxide Dismutase and High Activity of Glutathione Reductase in Erythrocytes From Centenarians, Age and Ageing, 1998