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It is a tetrameric protein with each subunit divided into four domains, the N-terminal threading arm, C-terminal helices, wrapping loop, and barrel [20] (Figure 4)

Barrier failure mechanisms arise from convergent cell death pathways: necrosis (with release of HMGB1 and ATP), NOX2-mediated apoptosis, NLRP3/gasdermin D-dependent pyroptosis, and ferroptosis driven by glutathione peroxidase 4 deficiency

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