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In mice, smoke effects on villi and TJs integrity were associated with increased small intestine permeability [57] Nicotine plays a protective role in ulcerative colitis (UC) by modulating cholinergic pathways in immune cells and reducing their activation and cytokine expression [58], while in Chrons disease (CD) it increases intestinal permeability by interacting with claudin gene expression [59], leading to apoptosis in the follicle-associated epithelium [60] and change in the composition of the gut microbiota [61]

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doi: 10.1038/s41573-022-00390-x 141 Alcal-BarrazaSRLeeMSHansonLRMcDonaldAAFreyWH2ndMcLoonLK