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While FoxO1-sufficient CD4 + T cells give rise to effector T cells, pre-T FH , and GC T FH upon immunization, FoxO1 / CD4 + T cells generate pre-T FH cells with higher expression of T FH -defining markers (Bcl6, CXCR5, and PD-1) and lowered T cell zone chemokine receptors (CD62L, PSGL1) (119)

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