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Oxidative Stress The concept of oxidative stress has developed from the free radical theory of oxygen toxicity developed in the 1950s with studies such as those by Rebecca Gerschman showing that oxygen poisoning and X-irradiation can both produce oxygen-derived free radicals [35,36]

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Similarly, berberine alleviates ceramide-mediated renal fibrosis by modulating iron metabolism and oxidative stress (Wang et al., 2023)

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Mitochondria-targeted compounds like AP39 and AP123 also play roles in H 2 S regulation (Ger et al., 2016)