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Reduction of 3-ketosphinganine forms dihydrosphingosine, which is then typically N-acylated by one of six ceramide synthases (CerS1-CerS6) with 1426 carbon saturated or monounsaturated fatty acids, to form dihydroceramides, subsequently dehydrogenated to ceramides by dihydroceramide desaturase [111]

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